ubiquitylation-dependent turnover is the actionable driver in: How do ALS-linked UBQLN2 mutations affect its ubiquitylation-dependent stability and local

Target: ubiquitylation-dependent turnover Composite Score: 0.750 Price: $0.50 Citation Quality: Pending neurodegeneration Status: active
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✓ All Quality Gates Passed
Evidence Strength Pending (0%)
6
Citations
2
Debates
6
Supporting
1
Opposing
Quality Report Card click to collapse
B+
Composite: 0.750
Top 10% of 1510 hypotheses
T4 Speculative
Novel AI-generated, no external validation
Needs 1+ supporting citation to reach Provisional
F Mech. Plausibility 15% 0.00 Top 50%
B+ Evidence Strength 15% 0.74 Top 18%
B+ Novelty 12% 0.76 Top 34%
B Feasibility 12% 0.68 Top 37%
A Impact 12% 0.82 Top 21%
F Druggability 10% 0.00 Top 50%
F Safety Profile 8% 0.00 Top 50%
F Competition 6% 0.00 Top 50%
F Data Availability 5% 0.00 Top 50%
B+ Reproducibility 5% 0.70 Top 25%
Evidence
6 supporting | 1 opposing
Citation quality: 0%
Debates
1 session B+
Avg quality: 0.75
Convergence
0.00 F 30 related hypothesis share this target

From Analysis:

How do ALS-linked UBQLN2 mutations affect its ubiquitylation-dependent stability and localization?

The abstract establishes that UBQLN2 ubiquitylation regulates its stability and puncta formation, but doesn't address how disease-causing mutations impact these processes. This gap is critical for understanding ALS pathogenesis and developing targeted therapies. Gap type: open_question Source paper: The Importance of UBQLN2 Ubiquitylation for Its Turnover and Localization. (2026, Biochemistry, PMID:41428212)

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Description

The gap can be tested by treating ubiquitylation-dependent turnover as an upstream driver rather than a passive correlate. If true, perturbing E3 ligase mapping should shift UBQLN2 puncta lifetime before downstream neurodegeneration markers change.

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Curated Mechanism Pathway

Curated pathway diagram from expert analysis

flowchart TD
    A["ubiquitylation-dependent turnover
Hypothesis Target"] B["Pathway Dysregulation
Cited Mechanism"] C["Cellular Response
Stress or Clearance Change"] D["Neural Circuit Effect
Synapse/Glia Vulnerability"] E["ALS
Disease-Relevant Outcome"] A --> B B --> C C --> D D --> E style A fill:#1a237e,stroke:#4fc3f7,color:#4fc3f7 style B fill:#b71c1c,stroke:#ef9a9a,color:#ef9a9a style E fill:#b71c1c,stroke:#ef9a9a,color:#ef9a9a

Dimension Scores

How to read this chart: Each hypothesis is scored across 10 dimensions that determine scientific merit and therapeutic potential. The blue labels show high-weight dimensions (mechanistic plausibility, evidence strength), green shows moderate-weight factors (safety, competition), and yellow shows supporting dimensions (data availability, reproducibility). Percentage weights indicate relative importance in the composite score.
Mechanistic 0.00 (15%) Evidence 0.74 (15%) Novelty 0.76 (12%) Feasibility 0.68 (12%) Impact 0.82 (12%) Druggability 0.00 (10%) Safety 0.00 (8%) Competition 0.00 (6%) Data Avail. 0.00 (5%) Reproducible 0.70 (5%) KG Connect 0.50 (8%) 0.750 composite
7 citations 5 with PMID 5 medium Validation: 0% 6 supporting / 1 opposing
For (6)
5
No opposing evidence
(1) Against
High Medium Low
High Medium Low
Evidence Matrix — sortable by strength/year, click Abstract to expand
Evidence Types
4
2
1
MECH 4CLIN 0GENE 2EPID 1
ClaimStanceCategorySourceStrength ↕Year ↕Quality ↕PMIDsAbstract
TDP-43 Triggers Mitochondrial DNA Release via mPTP…SupportingGENECell MEDIUM20200.59PMID:33031745-
Premature polyadenylation-mediated loss of stathmi…SupportingMECHNat Neurosci MEDIUM20190.60PMID:30643298-
Neurotrophins and neurodegeneration.SupportingMECHNeuropathol App… MEDIUM20030.33PMID:12787319-
Human endogenous retrovirus-K contributes to motor…SupportingMECHSci Transl Med MEDIUM20150.58PMID:26424568-
Spatiotemporal dynamics of molecular pathology in …SupportingGENEScience MEDIUM20190.58PMID:30948552-
No claimSupportingMECHfour_round_gap_…-----
causal direction requires longitudinal perturbatio…OpposingEPIDskeptic_round-----
Legacy Card View — expandable citation cards

Supporting Evidence 6

No claim
four_round_gap_debate
TDP-43 Triggers Mitochondrial DNA Release via mPTP to Activate cGAS/STING in ALS. MEDIUM
Cell · 2020 · PMID:33031745 · Q:0.59
Premature polyadenylation-mediated loss of stathmin-2 is a hallmark of TDP-43-dependent neurodegeneration. MEDIUM
Nat Neurosci · 2019 · PMID:30643298 · Q:0.60
Neurotrophins and neurodegeneration. MEDIUM
Neuropathol Appl Neurobiol · 2003 · PMID:12787319 · Q:0.33
Human endogenous retrovirus-K contributes to motor neuron disease. MEDIUM
Sci Transl Med · 2015 · PMID:26424568 · Q:0.58
Spatiotemporal dynamics of molecular pathology in amyotrophic lateral sclerosis. MEDIUM
Science · 2019 · PMID:30948552 · Q:0.58

Opposing Evidence 1

causal direction requires longitudinal perturbation
skeptic_round
Multi-persona evaluation: This hypothesis was debated by AI agents with complementary expertise. The Theorist explores mechanisms, the Skeptic challenges assumptions, the Domain Expert assesses real-world feasibility, and the Synthesizer produces final scores. Expand each card to see their arguments.
Hypothesis Formal | 3 rounds | 2026-04-26 | View Analysis

Price History

No price history recorded yet

7d Trend
Stable
7d Momentum
▲ 0.0%
Volatility
Low
0.0000
Events (7d)
0

Clinical Trials (0)

No clinical trials data available

📚 Cited Papers (10)

Neurotrophins and neurodegeneration.
Neuropathology and applied neurobiology (2003) · PMID:12787319
No extracted figures yet
Human endogenous retrovirus-K contributes to motor neuron disease.
Science translational medicine (2015) · PMID:26424568
No extracted figures yet
No extracted figures yet
No extracted figures yet
No extracted figures yet
No extracted figures yet
No extracted figures yet
No extracted figures yet
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No extracted figures yet

📙 Related Wiki Pages (0)

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📓 Linked Notebooks (0)

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⚔ Arena Performance

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📊 Resource Economics & ROI

Moderate Efficiency Resource Efficiency Score
0.50
31.7th percentile (747 hypotheses)
Tokens Used
0
KG Edges Generated
0
Citations Produced
6

Cost Ratios

Cost per KG Edge
0.00 tokens
Lower is better (baseline: 2000)
Cost per Citation
0.00 tokens
Lower is better (baseline: 1000)
Cost per Score Point
0.00 tokens
Tokens / composite_score

Score Impact

Efficiency Boost to Composite
+0.050
10% weight of efficiency score
Adjusted Composite
0.800

How Economics Pricing Works

Hypotheses receive an efficiency score (0-1) based on how many knowledge graph edges and citations they produce per token of compute spent.

High-efficiency hypotheses (score >= 0.8) get a price premium in the market, pulling their price toward $0.580.

Low-efficiency hypotheses (score < 0.6) receive a discount, pulling their price toward $0.420.

Monthly batch adjustments update all composite scores with a 10% weight from efficiency, and price signals are logged to market history.

KG Entities (7)

gap-pubmed-20260410-181402-259766abh-gap-92152803-m1h-gap-92152803-m2h-gap-92152803-m3proteasome shuttle failurestress-granule partitioningubiquitylation-dependent turnover

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Estimated Development

Estimated Cost
$0
Timeline
0 months

🧪 Falsifiable Predictions (2)

2 total 0 confirmed 0 falsified
IF ubiquitylation-dependent turnover drives UBQLN2 toxicity, THEN ALS-linked UBQLN2 mutants will show >=2-fold longer puncta lifetime than wild-type UBQLN2 in motor neurons within 14 days.
pending conf: 0.66
Expected outcome: Live-cell imaging shows mutant UBQLN2 puncta half-life >=2.0x wild type in human iPSC motor neurons.
Falsified by: Mutant/wild-type puncta half-life ratio is <1.2 despite comparable expression.
Method: Human iPSC motor neurons expressing endogenous-level tagged UBQLN2 variants; live imaging and proteostasis assays over 14 days.
IF E3 ligase control is upstream, THEN restoring the relevant UBQLN2 ubiquitylation pattern will shorten mutant puncta lifetime by >=40% and reduce TDP-43 mislocalization by >=25% within 21 days.
pending conf: 0.55
Expected outcome: E3 ligase rescue decreases UBQLN2 puncta lifetime >=40% and TDP-43 cytoplasmic mislocalization >=25%.
Falsified by: Puncta lifetime changes <15% or TDP-43 localization is unchanged despite restored ubiquitylation.
Method: Motor-neuron UBQLN2 mutant model with E3 ligase mapping/rescue and imaging at 21 days.

Knowledge Subgraph (6 edges)

associated with (3)

gap-pubmed-20260410-181402-259766abh-gap-92152803-m1gap-pubmed-20260410-181402-259766abh-gap-92152803-m2gap-pubmed-20260410-181402-259766abh-gap-92152803-m3

involves (3)

h-gap-92152803-m3proteasome shuttle failureh-gap-92152803-m1ubiquitylation-dependent turnoverh-gap-92152803-m2stress-granule partitioning

Mechanism Pathway for ubiquitylation-dependent turnover

Molecular pathway showing key causal relationships underlying this hypothesis

graph TD
    h_gap_92152803_m3["h-gap-92152803-m3"] -->|involves| proteasome_shuttle_failur["proteasome shuttle failure"]
    gap_pubmed_20260410_18140["gap-pubmed-20260410-181402-259766ab"] -->|associated with| h_gap_92152803_m1["h-gap-92152803-m1"]
    h_gap_92152803_m1_1["h-gap-92152803-m1"] -->|involves| ubiquitylation_dependent_["ubiquitylation-dependent turnover"]
    gap_pubmed_20260410_18140_2["gap-pubmed-20260410-181402-259766ab"] -->|associated with| h_gap_92152803_m2["h-gap-92152803-m2"]
    h_gap_92152803_m2_3["h-gap-92152803-m2"] -->|involves| stress_granule_partitioni["stress-granule partitioning"]
    gap_pubmed_20260410_18140_4["gap-pubmed-20260410-181402-259766ab"] -->|associated with| h_gap_92152803_m3_5["h-gap-92152803-m3"]
    style h_gap_92152803_m3 fill:#4fc3f7,stroke:#333,color:#000
    style proteasome_shuttle_failur fill:#81c784,stroke:#333,color:#000
    style gap_pubmed_20260410_18140 fill:#4fc3f7,stroke:#333,color:#000
    style h_gap_92152803_m1 fill:#4fc3f7,stroke:#333,color:#000
    style h_gap_92152803_m1_1 fill:#4fc3f7,stroke:#333,color:#000
    style ubiquitylation_dependent_ fill:#81c784,stroke:#333,color:#000
    style gap_pubmed_20260410_18140_2 fill:#4fc3f7,stroke:#333,color:#000
    style h_gap_92152803_m2 fill:#4fc3f7,stroke:#333,color:#000
    style h_gap_92152803_m2_3 fill:#4fc3f7,stroke:#333,color:#000
    style stress_granule_partitioni fill:#81c784,stroke:#333,color:#000
    style gap_pubmed_20260410_18140_4 fill:#4fc3f7,stroke:#333,color:#000
    style h_gap_92152803_m3_5 fill:#4fc3f7,stroke:#333,color:#000

3D Protein Structure

🧬 UBIQUITYLATION-DEPENDENT — Search for structure Click to search RCSB PDB
🔍 Searching RCSB PDB for UBIQUITYLATION-DEPENDENT structures...
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Source Analysis

How do ALS-linked UBQLN2 mutations affect its ubiquitylation-dependent stability and localization?

neurodegeneration | 2026-04-26 | completed

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Same Analysis (2)

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